The Daleks are an advanced warrior race of genetically-engineered mutants encased within tank-like shells that are bent on universal domination. To this end, the Daleks attempted to activate a reality bomb to eradicate all other forms of life in existence, but were stopped by an alliance of warriors from the Matoran Universe. Daleks view all other forms of life as inferior, but will ally themselves with other factions to suit their needs on occasion.
The Daleks rose to prominence when they created a reality bomb aboard the Dalek Crucible, which they intended to use to destroy the entire BZPB Multiverse and become the only forms of life in existence. Blackout became aware of this threat in 100,0XX CMT and formed a massive army that included Artrix, BoM Master, Kakamu, Malygos, TMV, Zev Raregroove, and countless others opposed to the Daleks' plan. The allies followed a Dalek saucer that had captured Findor back to the Dalek Crucible, but the Daleks destroyed much of their army with a single cannon blast and took the survivors onto the Crucible as prisoners. Blackout and Zev were both brainwashed and forced to attack their allies aboard the station.
Meanwhile, the Blackout, Overlord Blackout, Ynot, and Hollow Ynot of 150,080 CMT were transported back in time onto the Dalek Crucible through a crack in time. Ynot remembered that he did not participate in the battle against the Daleks when it happened, so he decided to hide to avoid the possibility that his presence would alter history. Ynot climbed up a large building to watch the battle from above, but the Daleks detected him and trapped him in a forcefield. Blackout then informed Ynot that Dalek forcefields "let stuff in, but not out", so Ynot looked for another means of escape and realized that he owned a Memory Seal that could return him to his own time. Ynot activated the Memory Seal and returned to 150,080 CMT, but the Blackout, Overlord Blackout, and Hollow Ynot of that time were transported with him because they were all linked to the same crack in time.
Afterwards, Malygos teleported to the main control room of the Crucible and deactivated the Dalek Supreme, which broke the Dalek's influence over Blackout and Zev's minds. Malygos then activated the Crucible's self-destruct sequence to neutralize the Dalek threat. The survivors that the Daleks had captured were teleported out as the Crucible and its Dalek inhabitants exploded behind them.
In 150,080 CMT, the Daleks returned in the Battle of Lantea and attacked Atlantis. The Daleks destroyed Axel Storm's vehicle, the Typhoon, but Blackout stole Atlantis and escaped with it intact before the Daleks could disable the city-ship.
In the Battle of BZPower, Blackout used the Architect's computer to create an amalgam fleet of thousands of alien ships that Blackout had programmed to be loyal to him regardless of their natural affiliations. D-Klak noted that part of the fleet consisted of Dalek ships. Blackout then ordered the fleet to attack his enemies below, which kicked off a massive battle. Dead Daleks were observed to fall around D-Klak as he plummeted towards the surface and landed, then teamed up with the BLUE Squad to attack more Dalek ships.