Starscream

Starscream was created 149,450 years ago in order to drive the Vrai out of the Matoran Universe along with Blackout, Barricade, Brawl and Bonecrusher. After this was accomplished, Starscream, Barricade, Brawl and Bonecrusher were placed in stasis pods. When Blackout created Darkmount, he took the pods with him, and later ordered Malygos to retrieve a crystal, hidden by the Great Beings on a distant planet, which would reawaken the Makuta. That happened. Then, Ynot, the Makuta Anima and The Fallen attacked. Starscream attacked Trixitin.

Later, Blackout summoned him and Grindor through a portal to fight Kakamu. He got hit with a Sleep beam. Then Malygos stole his body. Then he betrayed Blackout and took over the Twilight, which caused everyone to fight him.

During the Battle of Mata Nui II, he fought Catalyx and Ravage.

He then got sucked into a plothole, being revived by Xavalis and fighting alongside Blackout at the same time. He commanded Darkmount for a while during the Battle of Lantea, until Blackout came back from Atlantis.

Then Blackout sacrificed him to open the Pandorica. His armour was destroyed, but his antidermis survived, and travelled to BZPower, where he got a new body.....only to be captured by Sideways and used to revive Unicron.