Starscream

Starscream was created 149,450 years ago in order to drive the Vrai out of the Matoran Universe along with Blackout, Barricade, Brawl and Bonecrusher. After this was accomplished, Starscream, Barricade, Brawl and Bonecrusher were placed in stasis pods. When Blackout created Darkmount, he took the pods with him, and later ordered Malygos to retrieve a crystal, hidden by the Great Beings on a distant planet, which would reawaken the Makuta. That happened. Then, Ynot, the Makuta Anima and The Fallen attacked. Starscream attacked Trixitin.

Later, Blackout summoned him and Grindor through a portal to fight Kakamu.

He then took over the Twilight and betrayed Blackout, which caused everyone to fight him.

He then got sucked up into a plothole, being revived by Xavalis and fighting alongside Blackout at the same time. He commanded Darkmount for a while during the Battle of Lantea, until Blackout came back.